(de Haan, 1835)
Carapace covered by large granules, small central area totally devoid of granules; front deflexed, constricted between bases of ocular peduncles, with smooth margins, clearly bilobed distally, median furrow deep; lateral margins posteriorly slightly convergent, 2 well defined and 1 faint antero-lateral teeth. Ocular peduncles long and narrow, cornea extending to base of external orbital angle. Central region of epistome excavated. Merus of third maxilliped markedly smaller than ischium. Palm of male cheliped elongate, outer face with row of large tubercles at upper margin, otherwise finely granular, inner face heavily granular; immovable finger deflexed, cutting edge proximally with a large, wedge-shaped, crenulated tooth, distal half with few large and numerous small granules; cutting edge of dactylus proximally with a large quadrangular, crenulated tooth, distally with a row of tubercular granules. Meri of ambulatory legs with few hairs; anterior face of carpus and propodus of third pereiopod naked. Tip of horny endpiece of male first pleopod truncate. (Türkay 1995)
Type locality: Japan.
Range: Japan - (de Haan, 1835, Tesch, 1915), Tokyo Bay and Nagasaki (Ortmann, 1894a), Mutsu Bay (Yokoya, 1928), Shimoda (Sakai, 1935), Asamushi, Shimoda, Ise Bay, Wakayama and Nagasaki (Sakai, 1939), Nishinotsu, Okinohata, Yanagawa City (Miyake, 1961), Sagami Bay (Sakai, 1965b), from Aomori Bay to Kyushu and Amami-Oshima (Sakai, 1976a), Amakusa (Yamaguchi et al., 1976), Nanki Shirahama (Miyake, 1983), Tokyo Bay, Shirahama, Wakayama Prefecture, Yoshino-gawa, Tokushima Prefecture, and Tatara-gawa, Fukuoka Prefecture (Wada & Sakai, 1989); Korea - (Kamita, 1941), Jinsen (Sakai, 1939), Hae-undae, Moseulpo. Namweon, Seogwipo, Gapa Island, Yeosu, Daeheugsan Island, Bi-geum Island, Mogpo, Daecheon, Inch'on, Jag-yag island, Palmi Island, Seondu Ri, Seopo Ri, Yeong-jong Island, and Baegryeong Island (Kim, 1973), Kusan, Anmyun Island and Seoul (Wada & Sakai, 1989); China - Dairen (Sakai, 1939), Qingdao, Shandong (Wada & Sakai, 1989), Liaodong Peninsula, Liaodong Bay, Bo Hai, and Shandong Peninsula (Shen, 1932), Hainan Island (Shen, 1936), Ningbo, Tsimei, Xiamen, and Hainan Island (Shen, 1940b), Habei, Haimen, Quangdang, Beidaihe, and Yanmadao (Tai & Song, 1984), Hainan Island, Fujian, and Zhejiang (Dai & Yang, 1991); Taiwan - Tan-shui (Sakai, 1939), Lu-kang, Peimunyu, Chiayi, and Tan-shui (Lin, 1949), Dan-Shui River, Taipei County, Wu Creek, Taichung County, San-Tiou-Lon, Yunlin County, Buh-Dai, Jiayi County, and Chi-Guh, Tainan County (Huang et al., 1992); Singapore (Barnes, 1970); Australia - Shark Bay (Barnes, 1967); tidal muddy flats or river mouth.