Takeda, 1971
Diagnosis: Small species. Carapace transversely ovate and evenly convex in both directions. Dorsum with granulated areolae. Front with two lobes double-rimmed and fringed with two rows of pearly granules. Antero-lateral border with four conical teeth behind external orbital angle. Chelipeds heavy with abruptly curved and gaped fingers.
Description of holotype: The carapace is more or less transversely ovate and only evenly convex in both directions; the dorsal surface is well separated into regions which are not strongly convex; the frontal region and the wide shallow furrow along the supraorbital and antero-lateral borders are rather sparsely covered with small warty granules, while most of the regions are roughened by numbers of transverse short ridges that are more or less beaded to form transverse indistinct rows; the short ridges are almost indistinct or absent on the posterior part of the mesogastric region (3M) and the cardiac region (1P); the areola 1M is more or less convex anteriorly and shallowly separated from the inner part of 2M which is imperfectly subdivided into two by a longitudinal shallow but distinct furrow on the anterior half; the outer part with a small tuft of feathered hairs on the anterior margin is about one and a half times the breadth of the inner part; in the median line 3M bears a pair of transverse slit-like furrow at the posterior end; the areola 4M is small but very distinct, being separated from 3M by a narrow deep furrow and from 1P by a wide deep depression; the areola 1L is only a small prominence obliquely behind and inside the first antero-lateral tooth; the areolae 2 and 3L are almost separated by an oblique furrow; the areola 3L is again more or less subdivided into two by a shallow transverse furrow, the anterior subregion being smaller and subequal to 2L; the areola 4L is convex and subequal to or only slightly larger than 2L, being well isolated from the adjacent regions and the antero-lateral tooth; the areola 5L is very large with a small tuft of feathered hairs near the middle of the anterior margin and imperfectly separated from 6L by the oblique anterior and posterior incisions; the areola 1P is also prominent and weakly convex towards the posterior border of the carapace in the median line; the areola 2P is rather strongly convex fore and aft near the lateral end just above the base of the last ambulatory leg; the postero-lateral dorsal surface is ornamented with a furrow which is extended towards the posterior angle of 6L.
The front is rather prominent and cut into two lobes by a median deep notch, being distinctly double-rimmed with fringes of two rows of pearly granules; each lobe is more extended forwards near the median notch and weakly retreats laterally, bearing a rounded, rather strong lateral lobule. The supraorbital border bears two notches as usual and fringed with a row of granules. The infraorbital border is rather deeply concave, the inner infraorbital angle being strongly extended forwards and subacute at the apex. The antero-external angle of the antennal basal segment is a little prolonged into the orbit, but imperfectly shuts out the flagellum from the orbit.
The antero-lateral border of the carapace is armed with four teeth excluding the external orbital angle which is conical with an acute apex and rather similar to the antero-lateral teeth; the four antero-lateral teeth are similar to each other, conical and fringed with granules, the first being the smallest of the series; in each of the first and third the posterior slope is slightly longer than the anterior; in all the teeth the granules at the ends of the slopes are larger, but rounded and not protruded from the general contour of the border; each of the third and fourth bears a longish seta at the dorsal part.
The chelipeds are heavy, the right being only slightly larger than the left. The merus is granulated and exerted beyond the carapace by its distal part; the anterior and upper borders are fringed with longish feathered hairs; the anterior border is bluntly crested and bears a row of granules, some of which near the proximal part are prominent, while the upper border is rounded. The outer surface of the carpus is rather reticulated with scattered and beaded granules and bears a rounded prominence near the articulation with the palm; the inner angle is armed with two spiniform teeth, the upper of which is much the larger and curved upwards. The palm is very high and the oblique rugosities with the ridges of low fused granules on the outer surface are everywhere prominent; in the smaller chela the ornamentation mentioned is more prominent than in the larger chela. The immovable finger bears a deep depression with the granulated ridge along the lower border and a granulated ridge along the prehensile edge; the lower borde is rather strongly convex in the middle; in both chelae each movable finger is abruptly curved to be nearly vertical near the tip, leaving a wide gape between both fingers; in the larger chela the prehensile edge of the immovable finger bears altogether four teeth at the proximal half, the distal one of which is very prominent and high, and the movable finger is provided with two, the proximal one of which is molar-like and the distal one is conical; the formation of the smaller chela is rather similar to that of the larger, but the teeth are much smaller and conical, especially the proximal two of the immovable finger are very small and granuliform; each of the outer surfaces of all the fingers is provided with two more or less distinct hairs in the middle near the prehensile edge; the distal halves of the fingers are deeply hollowed and provided with a prominent tuft of hairs in the immovable finger and with a prominent and two small tufts in the movable finger; the colour of the immovable finger is not much extended back onto the palm.
The ambulatory legs are hairy and spinulated; The anterior border of the merus is fringed with feathered hairs and bears more than fifteen conical granules. The anterior border of the carpus and both borders of the propodus are thickly covered with conical granules of various size. The dactylus is also armed with conical granules, of which several on the anterior border and one on the subterminal part of the posterior border are spiniform.
In the abdomen, the right halves of the second and third segments are somewhat damaged. The first pleopod bears a very small beak only with some short subterminal hairs. (Takeda, 1971)
Type locality: Ngardmau, Babeldaob Island, Palau, 7°37'N, 134°33'E, 26 m.
Range: Kenya - Mombasa (Serène, 1984); Taiwan; Palau (Takeda, 1971).