Charybdis variegata

(Fabricius, 1798)

Carapace densely pubescent, granulate in postfrontal area and in the vicinity of the antero-lateral borders; transverse granular lines on protogastric and mesogastric regions, epibranchial line interrupted at the cervical groove and across midline, cardiac region with 1, each mesobranchial region with 3 such lines; front with 6 teeth, medians and submedians truncate, laterals triangular with rounded tips, overreaching the other teeth; antero-lateral borders with 6 teeth, first dentiform, last tooth spiniform, twice size of preceeding, second and third with rounded lateral borders and tips, fourth and fifth acute; postero-lateral junctions rounded. Antennal flagellum excluded from orbit. Chelipeds covered with large granules and/or squamiform markings; merus with 3 strong spines and numerous granules on anterior border, posterior border granulate; carpus with a strong internal spine, outer border with 3 spinules; palm with 4 spines on upper border, lower border longitudinally sulcate. Merus of swimming leg with a subdistal posterior spine, propodus with 1-2 of denticles on posterior border. (Türkay 1995)

Antennal flagellum excluded from orbital hiatus; median and submedian frontal teeth prominent beyond laterals, median frontal teeth triangular, projecting anteriorly to level of inner infraorbital angle; posterior border of cephalothorax curved, forming curved postero-lateral junction; 6 antero-lateral teeth; second antero-lateral tooth not much smaller than first; distinct cardiac and mesobranchial ridges on carapace; lateral borders or penultimate segment of male abdomen convex; male first gonopod with lip membrane distinctly lobed and separated from tip. (Davie 1999)

Type locality: "Oceano Indico", probably Tranquebar, India (Apel & Spiridonov, 1998: 210).
Range: Red Sea - Aseb (Nobili, 1906); Persian Gulf (Alcock, 1899a); South Africa - off Tugela River mouth (Stebbing, 1920b, Barnard, 1950); Madagascar; Pakistan - Karachi (Apel & Spiridonov, 1998); India - Madras (Alcock, 1899a); West Malay Peninsula - Thailand (Stephenson, 1972a); Japan - Nagasaki (Alcock, 1899a), Kinkazan (Yokoya, 1933), Shimoda (Sakai, 1935), Shimoda, Ise Bay and Nagasaki (Sakai, 1939), Sagami Bay (Sakai, 1965b), Sagami Bay, Shimoda, Mikawa Bay, Kii Nagashima, Kii Minabe, and Nagasaki (Sakai, 1976a), Amakusa (Yamaguchi et al., 1976), Mikawa Isshiki and Toba (Muraoka, 1998); Taiwan - Taiwan Strait (Stephenson, 1972a), Kaohsiung and I-lan Provinces (Huang & Yu, 1997); China - Hong Kong (Alcock, 1899a), Tsimei and Xiamen (Shen, 1940b), Namoa (Stephenson, 1972a), Guangdong and Fujian (Dai & Yang, 1991); Philippines; Singapore (Shen, 1937); Indonesia - off Djangkar, Java (Leene, 1938); Australia - Northern Territory; 9-85 m.

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