(A. Milne Edwards, 1861)
Carapace covered with a dense short tomentum; transverse granular lines, protogastric and mesogastric regions, epibranchial line interrupted at the cervical groove and across midline; front with 6 acute triangular teeth, medians the broadest and most prominent, laterals the narrowest; antero-lateral borders with 6 acute dark tipped teeth, first with a very slightly concave lateral border, sixth narrowest; postero-lateral junctions rounded. Antennal flagellum excluded from orbit. Chelipeds densely hairy; merus with 3 strong spines on anterior border, posterior border smooth; carpus with a strong internal spine, outer border with 3 spinules; palm with 5 spines on upper border. Merus of swimming leg with a subdistal posterior spine, propodus smooth on posterior border. (Türkay 1995)
Antennal flagellum excluded from orbital hiatus; posterior border of cephalothorax curved, forming curved postero-lateral junction; 6 antero-lateral teeth, first antero-lateral tooth truncate; no distinct cardiac ridges on carapace; anterior border of arm of cheliped with 3 or more spines; hand of cheliped with 5 well developed spines; merus of fifth leg about 1.5 times as long as broad; penultimate segment of male abdomen with sides gradually converging. (Davie 1999)
Type locality: Japan.
Range: Japan - (de Haan, 1835, A. Milne Edwards, 1861, Parisi, 1916, Leene, 1938), Tokyo Bay and Maizuru (Ortmann, 1893), Tokyo Bay, Yokohama, Hondo, and Yokosuka (Parisi, 1916), Mutsu Bay (Yokoya, 1928), Shimoda (Sakai, 1935), Tateyama Bay, Manazuru, Shimoda, Ise Bay, Nanki Shirahama and Nagasaki (Sakai, 1939), Sagami Bay (Sakai, 1965b), Nagasaki (Stephenson, 1972a), from Tokyo Bay to Kyushu (Sakai, 1976a), Amakusa (Yamaguchi et al., 1976), Yamagata Prefecture (Suzuki S., 1979), Kushimoto (Miyake, 1983), Oshoro, Kikonai and Hakodate, Hokkaido (Komai et al., 1992), Okinawa; Korea - Kankyo-hokudo, Kankyo-nando, Kogen-do, Keisho-hokudo, Keisho-nando, Zenra-nando, Zenra-hokudo, Chusei-nando, Keiki-do, Kokai-do and Heian-nando (Kamita, 1941), Pohang, Yeong-jong Island, Deogjeog Island, Ongdo Island, Anmyeon Island, Jag-yag Island, Daecheong Island, Daeyeonpyeong Island, Gomso, Bi-geum Island, Yeosu, Hae-undae, Jinhae, Sangchuja Island, Seogwipo, and Seongsanpo (Kim, 1973); Taiwan - Tan-shui (Sakai, 1939), Hsin-chu, Chi-lung, and Tan-shui (Lin, 1949), Taiwan Strait (Stephenson, 1972a), T'ai-chung, Yüan-lin and Kaohsiung Provinces (Huang & Yu, 1997); China - Fuzhou (Rathbun, 1931), Liaodong Peninsula, Liaodong Bay, Bo Hai Gulf, and Shandong Peninsula (Shen, 1932), Xiamen (Leene, 1938), Shenkiamen, Fuzhou, Liuwutien, Tsimei, and Xiamen (Shen, 1940b), Guangdong, Fujian, Zhejiang, Shandong Peninsula, and Liaodong Peninsula (Dai & Yang, 1991); Hawaiian Islands - Honolulu (Rathbun, 1906, this record seems to be an error; Sakai, 1976: 355): littoral to 15 m.