(Takeda, 1995)
Description of holotype: Carapace nearly as long as wide, strongly convex dorsally for its most part behind fronto-orbital region, armed with 16 dorsal and 9 marginal tubercles; dorsal surface and margin of carapace seemingly smooth, though thickly spaced with microscopical granules; most of dorsal and marginal tubercles conical, but somewhat different in size and shape; protogastric main tubercle stout, too sharp to be regarded as a mound, but too blunt to be called a tubercle; protogastric subsidiary tubercle much smaller than, and situated obliquely in front of the main one; mesogastric region indicated only by a weak longitudinal ridge; cardiac tubercle as large as the main protogastric one, but distinctly tuberculated; of 6 branchial tubercles, anterior 2 of outer 3 tubercles and both of inner 2 tubercles conical, distinctly tuberculated, similar in size and shape to cardiac tubercle; anterior one of inner 2 tubercles just in a transverse line with main protogastric tubercle, and posterior one in a transverse line with intestinal tubercle; posteriormost of outer 3 tubercles markedly developed, strongest of all the dorsal tubercles, more than twice as long as other dorsal tubercles, obliquely directed posteriorly as a whole, curved dorsally near its tip; intestinal tubercle rather close to posterior border of carapace, similar to, but slightly shorter than main branchial tubercle, more or less depressed in dorsal view, directed posteriorly, with blunt apex.
Of 9 marginal tubercles of carapace, hepatic and branchial ones nearly the same in size, the former directed obliquely forwards just like anterior branchial one, but much sharper; median one of 3 tubercles on posterior margin slightly smaller than lateral ones.
Supraorbital tubercle bluntly tuberculated, with convex dorsal surface; frontal region between supraorbital tubercles of both sides deeply excavated, somewhat eroded near its anterior margin, with a small median U-shaped incision.
Two long fused slits on supraorbital margin, and 1 on infraorbital margin; carapacial margin following external orbital angle rather thickened and directed towards anterior branchial tubercle. True carapacial margin directed towards antero-external angle of buccal cavern, which forms a part of roof and wall of distal end of spout and extends beyond fronto-orbital margin in dorsal view.
Third maxilliped long, roughened with tubercles mainly on merus and distal third of exopod, being studded with several pearly granules in a longitudinal line on median part of ischium and along outer margin of exopod; merus half as long as ischium, and exopod slightly narrower than ischium.
Chelipeds slender, mostly cylindrical, thickly covered with microscopical granules; merus only slightly longer than carpus and palm combined; palm slightly longer than fingers, distinctly tapering; both fingers weakly sinuate, leaving a narrow gape between them for their distal halves to two-thirds, with comb-like teeth armed with many perpendicular needle.
All of chelipeds and ambulatory legs detached. Ambulatory legs cylindrical and microscopically granulated, with thick setae on dactyli.
Proximal two abdominal segments linear; main fused segment long, convex ventrally at each side of basal third; distal two-thirds of main segment flattened for its most part, only with a small thin tubercle just near distal margin, with lateral margins of both sides subparallel or only weakly tapering distally; terminal segment more or less quadrate.
First pleopod curved outwards to be sickle-shaped at distal third, with a distal large flap of right-angled triangle frilled with spinules along hypotenuse. (Takeda, 1995)
Type locality: Ie-jima Islet, west of Okinawa-jima, Japan, in front of Large Cave, 26°42.9'N, 127°50.1'E, 35 m.
Range: Japan - Ie-jima Islet, west of Okinawa-jima (Takeda, 1995); Philippines, Indonesia, New Caledonia, Chesterfield Island, Madagascar (Galil, 2001c); 35 m.