Crosnier, 1962
Carapace very broad (breadth about 2 times length); surface with elevated granular areas; regions well recognizable; front with 4 low set and rounded lobes, outer pair very much broader and longer than inner ones; antero-lateral borders with 6-9 (mostly 8-9) teeth, last one very large and projecting obliquely backwards; postero-lateral junction elevated and bearing a spine. Antero-external angle of merus of third maxillipeds rounded, not markedly produced into a lobe. Chelipeds moderately long (not more than 2.5 times as long as carapace), merus with 1 spine at postero-distal border, anterior border with 3-4 spines; carpus with 2 spines; upper surface of palm with 2 distal spines, lower surface smooth to naked eye. Posterior border of merus of swimming leg very finely serrated, remaining joints smooth. (Türkay 1995)
Postero-lateral junction of carapace angular or spinous; front distinctly 4-toothed or 4-lobed, median frontal teeth very much shorter than, and less prominent than, laterals; lateral frontal teeth triangular, projecting, and typically sharp; carapace moderately strongly embossed, with tuberculate but not spiniform elevations in cardiac and mesobranchial areas; metagastric area with short ridges, and without tubercles; posterior border of arm of cheliped with single spine; upper surface of palm of cheliped with 2 distal spines, or 1 spine and 1 tubercle; chelipeds of moderate length and robustness, less than 2.5 times the carapace length. (Davie 1999)
Type locality: Nosy Iranja, Madagascar, intertidal.
Range: Madagascar - Nosy Iranja (Crosnier, 1962); Japan -Yoron-jima, Amami-shoto (Sakai, 1976a); Taiwan - Nan-Wan and Wanlitung, P'ing-tung Province (Huang & Yu, 1997); China - Paracel Islands (Chen H., 1980b, Dai & Yang, 1991); Sulu Archipelago - Tawitawi (Stephenson, 1975); shallow waters.