Chen, 1980
Carapace very strongly sculptured, with deep grooves and conspicuous wart-like tubercles; antero-lateral margin, between base of exorbital tooth and cervical groove, smooth, without tubercles or denticles. Teeth on lower orbital margin in cluster. Carpus of cheliped distinctly granulated on upper surface and with a conspicuous row of granules along anterior margin. Merus of second and third pereiopods almost cylindrical. (Türkay 1995)
Type locality: Nan'ao Island, Guangdong Province, China.
Range: Japan - (de Haan, 1841, White, 1847f, Herklots, 1861, Miers, 1884a, Sakai, 1930, 1936, 1956, 1980, Utinomi, 1956, 1969, Nishimura & Suzuki, 1971, Takeda, 1975b, 1982, 1983), Tokyo Bay, Kochi and Kagoshima (Ortmann, 1892), Sagami Bay (Doflein, 1902, Parisi, 1914), Wakanoura Bay (Rathbun, 1902a), Nagasaki (Rathbun, 1920, Sakai, 1934a), Boso Peninsula, Ito, Sagami Bay and Nagasaki (Balss, 1922b), Kagoshima (Urita, 1926a), Tanabe Bay (Komai et al., 1927), Shimoda (Sakai, 1935a), Tateyama Bay (Sakai, 1936b), Tokyo Bay, Sagami Bay, Shimoda, Izu Peninsula, Ise Bay, Kii Peninsula, Tosa Bay, Nagasaki and Kagoshima (Sakai, 1937a), Tokyo Bay (Sakai, 1940), Mie Prefecture (Shiino & Yamada, 1951), Amakusa (Miyake, 1961b, Yamaguchi et al., 1976, Takeda, 1978a, Yamaguchi et al., 1987), Tsuyazaki, Amakusa, Sea of Ariake, Tottori, Toyama, and Yamagata (Miyake et al., 1962), Sagami Bay (Sakai, 1965b), Tokyo Bay, Sagami Bay and Oga Peninsula (Sakai, 1976a), Yamagata Prefecture (Suzuki S., 1979), Enshu-nada (Takeda, 1981), Izu Peninsula (Takeda, 1982a), Kii Minabe and Tokushima Prefecture (Sakai K. & Nakano, 1983), Tosa Bay (Miyake, 1983, Quintana, 1987), Hakata Bay (Hamano et al., 1985), Sagami Bay, Tanabe Bay, Tosa Bay, Nagasaki and probably near Nagasaki, Amakusa, Suruga Bay and near Kobe (Holthuis & Manning, 1990); China - (?Berthold, 1845, ?Neuville, 1938), Fujian, Fuzhou and Guangdong (Holthuis & Manning, 1990), Guangdong (Chen, 1980a), Hainan Island (Dai & Yang, 1991); Hong Kong (Holthuis & Manning, 1990); 15-118 m.