(de Haan, 1841)
Carapace as broad as long or slightly broader, flattened, all its surface coarsely granular; cervical and branchial grooves well marked. Chelipeds very unequal in larger males (carapace length > 15 mm); outer surface of palm and carpus granular, fingers smooth. Second and third pereiopods long, merus shorter than carapace, with merus and carpus coarsely granular, and merus, carpus, and propodus pubescent. (Türkay 1995)
Type locality: Japan
Range: Russia - Vladivostok (Balss, 1922b, Holthuis & Manning, 1990), Zaliv Petri Velikogo (Levin, 1976), Zaliv Posyete (Kobjakova, 1967, 1971, Vaskovsky & Suppes, 1972), off Kril'on, Sakhalin (Urita, 1942); Japan - (de Haan, 1841, Herklots, 1861, Bouvier, 1899, Parisi, 1914, Sakai, 1930, 1936, 1940, 1956, 1980, 1985, Shimoizumi & Tanemura, 1950, Nishimura & Suzuki, 1971, Takeda, 1975b, 1982, 1983, Muraoka, 1982, Miyake, 1983, Sakai et al., 1983, Holthuis & Manning, 1990), off north-eastern coast of Japan and Hakodate (Stimpson, 1858c, 1907), Yokohama (Targioni Tozzetti, 1872b, 1877, Nobili, 1903c), Sagami Bay (Döderlein, 1883), Katsuyama, Tanagawa, Tokyo Bay, and Kagoshima (Ortmann, 1892), Miyako and Wakanoura (Rathbun, 1902a), Ishikawa Prefecture (Anonymous, 1904), Tokyo Bay, Yokohama and Suruga Bay (Parisi, 1914), Aomori, Onagawa Bay, Kominato, Tokyo Bay, Sagami Bay, Ise Bay, Osaka Bay, and Nagasaki (Balss, 1922b), Kagoshima (Urita, 1926a, 1942), Niigata and Toyama Bay (Kamita, 1933), Kinkazan, Todo-saki, between Siwoya-zaki and Kinkazan, Suruga Bay, Inubo-zaki, Owase, Goto-retto, Tsushima Strait, Oga Peninsula and west of Jusan-gata (Yokoya, 1933), Nagasaki (Sakai, 1934a), Shimoda (Sakai, 1935a), Tokyo Bay, Sagami Bay, Ise Bay and Nagasaki (Sakai, 1937a), Otaru, Shioya, Toyama Bay, Katsuyama, and Sagami Bay (Urita, 1942), Mie Prefecture (Shiino & Yamada, 1951), Oshoro Bay (Uchida T., 1960), Amakusa and Sea of Ariake (Miyake, 1961a, b), Amakusa, Sea of Ariake, Tsuyazaki, Toyama, and Niigata (Miyake et al., 1962), San-in District (Kamita, 1963), Hokkaido (Kurata, 1964), Sagami Bay (Sakai, 1965b), Mikawa Bay (Serène & Romimohtarto, 1969, Yang C.M., 1979), off Tsushima (Takeda, 1973), Mutsu Bay (Takeda, 1974b), Sagami Bay, Mikawa Bay, Kii Nagashima, and Kii Minabe (Sakai, 1976a), Aitsu (Yamaguchi et al., 1976), Amakusa (Takeda, 1978a, Yamaguchi et al., 1987), Yamagata Prefecture (Suzuki S., 1979), Enshu-nada (Terada, 1981), Todo-saki, between Siwoya-zaki and Kinkazan, Inubo-zaki, Owase, Goto-retto, Tsushima Strait, Oga Peninsula and west of Lake Zyusan-gata (Horikoshi et al., 1982), Sagami Bay (Takeda, 1982), Suruga Bay (Horikoshi et al., 1983), Tanabe Bay (Miyake, 1983), Tokushima Prefecture (Sakai K. & Nakano, 1983), Hakata Bay (Hamano et al., 1985), Tosa Bay (Quintana, 1987), Otaru, Hakodate, Kinkazan, Yamagata Prefecture, Toyama Bay, Miyako, Tokyo Bay, Sagami Bay, Kii Peninsula, Tanabe Bay, Osaka Bay, Awajishima, Tosa Bay, Nagasaki, and Amakusa (Holthuis & Manning, 1990), Soya Strait (Takeda & Miyauchi, 1992), Hakodate, Oshoro, Rishiri and Utoro (Komai et al., 1992), off Matsuyama, Seto Inland Sea (Takeda et al., 2000); Korea - (Kamita, 1934, 1935, 1936), Kankyo-hokudo, Kankyo-nando, Kogen-do, Keisho-nando, Zenra-nando and Chusei-nando (Kamita, 1941b), Ju-munjin, Bangjugpo, Jinhae, Namcheon Dong, Yeong-il Bay, Hae-undae, and Sogcho (Kim, 1973); East China Sea (Takeda & Miyake, 1970a, 1972c); China - (Yang S., 1986, Chen H., 1986b), north China (Shen, 1937), Chinese Seas (Gee, 1925), Liaoning Province (Zhou & Sun, 1986), Penglai and Weihai (Shen, 1932b), Liuwutien, Jimei and Xiamen (Shen, 1940b), Jimei, Xiamen, Qingdao and Zhoushan, (Holthuis & Manning, 1990), Guangdong including Hainan Island (Dai & Yang, 1991), Hong Kong (Ortmann, 1892, Stimpson, 1858c, 1907, Shen, 1931a, 1940, Holthuis & Manning, 1990); Taiwan - off Chilung (Takeda & Miyake, 1970a), Tingch'ieting (Lin, 1949).