Literature
- Balss, H. 1922c. Östasiatische Decapoden. IV. Die Brachyrhynchen (Cancridea). Archiv für Naturgeschichte, 88A(11): 94-166, figs 1-2, pls 1-2.
- Sakai, T. 1939. Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha, pp. 365-741, figs 1-129, pls 42-111, table 1. Yokendo Co., Tokyo.
- Sakai, T. 1976a. Crabs of Japan and the Adjacent Seas. (In 3 volumes: (1) English text: i-xxix, 1-773, figs 1-379, (2) Plates volume: 1-16, pls 1-251, (3) Japanese text: 1-461, figs 1-2, 3 maps.) Kodansha Ltd, Tokyo.
- Stephenson, W. 1972b. An annotated check list and key to the Indo-West-Pacific swimming crabs (Crustacea: Decapoda: Portunidae). Bulletin of the Royal Society of New Zealand, 10: 1-64.