Literature
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- Dana, J.D. 1852c. Crustacea. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U.S.N., 13(pt. 1): i-viii, 1-685, pl. 8.
- Edmondson, C.H. 1959. Hawaiian Grapsidae. Occasional Papers of the Bernice P. Bishop Museum, Honolulu, 22(10): 153-202, figs 1-27.
- Rathbun, M.J. 1918a. The Grapsoid Crabs of America. United States national Museum Bulletin, 97: i-xxii, 1-461, figs 1-72, pls 1-161.
- Rathbun, M.J. 1931. New and rare Chinese crabs. Lingnan Science Journal, 8 (1929): 75-104, pls 5-15.
- Sakai, T. 1939. Studies on the crabs of Japan. IV. Brachygnatha, Brachyrhyncha, pp. 365-741, figs 1-129, pls 42-111, table 1. Yokendo Co., Tokyo.
- Sakai, T. 1965b. The Crabs of Sagami Bay, collected by His Majesty the Emperor of Japan, i-xvi, 1-206 (English text), figs 1-27, pls 1-100: 1-92 (Japanese text): 1-26 (references and index in English): 27-32 (index in Japanese), 1 map. Maruzen Co., Tokyo.
- Sakai, T. 1976a. Crabs of Japan and the Adjacent Seas. (In 3 volumes: (1) English text: i-xxix, 1-773, figs 1-379, (2) Plates volume: 1-16, pls 1-251, (3) Japanese text: 1-461, figs 1-2, 3 maps.) Kodansha Ltd, Tokyo.
- Wada, K. 1995. Brachyura. In: S. Nishimura, Guide to seashore animals of Japan with color pictures and keys. Vol. 2: 379-418, pls 101-118. (In Japanese).